Raymond Grizzle

Phone: (603) 862-5130
Office: Biological Sciences, Jackson Lab, Durham, NH 03824
Ray Grizzle

Much of my recent research has involved restoration of oyster reefs and oyster aquaculture in New England, including integrated farming methods for oysters and seaweeds. My laboratory also has participated in a variety of seafloor mapping projects focusing on towed underwater video, but also including sonar and other remote sensing techniques. Other research topics include the effects of marine protected areas on seafloor ecology and fisheries, impacts of aquaculture activities on benthic communities, and succession of fouling communities on aquaculture cages.


  • Ph.D., Biology/Biological Sciences, Rutgers University
  • Ph.D., Ecology, Rutgers University
  • M.S., Biology, University of Central Florida
  • B.S., Biology, Florida State University

Courses Taught

  • INCO 590: Rsrch Exp/Biological Sciences

Selected Publications

Hoellein, T. J., Zarnoch, C. B., & Grizzle, R. E. (2015). Eastern oyster (Crassostrea virginica) filtration, biodeposition, and sediment nitrogen cycling at two oyster reefs with contrasting water quality in Great Bay Estuary (New Hampshire, USA). Biogeochemistry, 122(1), 113-129. doi:10.1007/s10533-014-0034-7

Grizzle, R. E., Ward, L. G., Fredriksson, D. W., Irish, J. D., Langan, R., Heinig, C. S., . . . Eberhardt, A. L. (2014). Long-term seafloor monitoring at an open ocean aquaculture site in the western Gulf of Maine, USA: Development of an adaptive protocol. Marine Pollution Bulletin, 88(1-2), 129-137. doi:10.1016/j.marpolbul.2014.09.014

zu Ermgassen, P. S. E., Spalding, M. D., Grizzle, R. E., & Brumbaugh, R. D. (2013). Quantifying the Loss of a Marine Ecosystem Service: Filtration by the Eastern Oyster in US Estuaries. Estuaries and Coasts, 36(1), 36-43. doi:10.1007/s12237-012-9559-y

Grizzle, R. E., Coen, L. D., & Rybak, A. (2011). TWO-DIMENSIONAL "ECOMAPS" OF SESTON DEPLETION ZONES OVER AN INTERTIDAL CRASSOSTREA VIRGINICA OYSTER REEF. JOURNAL OF SHELLFISH RESEARCH, 30(2), 512-513. Retrieved from http://gateway.webofknowledge.com/

Grizzle, R. E., Ward, L. G., Mayer, L. A., Malik, M. A., Cooper, A. B., Abeels, H. A., . . . Rosenberg, A. A. (2009). Effects of a large fishing closure on benthic communities in the western Gulf of Maine: recovery from the effects of gillnets and otter trawls. FISHERY BULLETIN, 107(3), 308-317. Retrieved from http://gateway.webofknowledge.com/

Grizzle, R. E., Greene, J. K., & Coen, L. D. (2008). Seston Removal by Natural and Constructed Intertidal Eastern Oyster (Crassostrea virginica) Reefs: A Comparison with Previous Laboratory Studies, and the Value of in situ Methods. Estuaries and Coasts, 31(6), 1208-1220. doi:10.1007/s12237-008-9098-8

Greene, J. K., & Grizzle, R. E. (2007). Successional development of fouling communities on open ocean aquaculture fish cages in the western Gulf of Maine, USA. Aquaculture, 262(2-4), 289-301. doi:10.1016/j.aquaculture.2006.11.003

Grizzle, R. E., Short, F. T., Newell, C. R., Hoven, H., & Kindblom, L. (1996). Hydrodynamically induced synchronous waving of seagrasses: ‘monami’ and its possible effects on larval mussel settlement. Journal of Experimental Marine Biology and Ecology, 206(1-2), 165-177. doi:10.1016/s0022-0981(96)02616-0

Grizzle, R. E., Langan, R., & Huntting Howell, W. (1992). Growth responses of suspension-feeding bivalve molluscs to changes in water flow: differences between siphonate and nonsiphonate taxa. Journal of Experimental Marine Biology and Ecology, 162(2), 213-228. doi:10.1016/0022-0981(92)90202-l

Grizzle, R. E., & Morin, P. J. (1989). Effect of tidal currents, seston, and bottom sediments on growth of Mercenaria mercenaria: results of a field experiment. Marine Biology, 102(1), 85-93. doi:10.1007/bf00391326

Most Cited Publications