Wilfred Wollheim

Co-director, Water Systems Analysis Group
Phone: (603) 862-5022
Office: Natural Resources & the Environment, James Hall Rm 276, Durham, NH 03824
Pronouns: He/him/his

In understanding the environment as an integrated system, WSAG explores the physical, chemical and biological processes that shape hydrological systems, with emphasis on the unique role of humans as agents of change. Our research and educational programs cross the boundaries of traditional scientific disciplines to foster interdisciplinary understanding of environmental change. Through field monitoring, whole ecosystem experiments, novel in situ sensor deployments, and dynamic hydrological and ecosystem modeling, we attempt to understand hydrological and biogeochemical dynamics at scales ranging from individual ecosystems, to whole river systems, to the global systems of inland waters. Built on strong collaborations that are national and international in scope, our group also serves as a teaching platform for the next generation of ecosystem and earth system scientists.

The success of the Water Systems Analysis Group reflects a rapidly emerging scientific field which pursues integrative studies of hydrology, biogeochemistry, and human-water interactions that is necessary to analyze the full dimension of anthropogenic change at local, regional, and global scales.


  • Ph.D., Earth Sciences, University of New Hampshire
  • M.S., Zoology/Physiology, University of Wyoming
  • B.S., Natural Resources, Cornell University

Courses Taught

  • NR 504: Freshwater Resources
  • NR 707/807: Environmental Modeling
  • NR 751/851: Aquatic Ecosystems
  • NR 791: Proj/Environmental Science I
  • NR 795W: Investigations
  • NR 905: Grant Writing
  • NR 993: Sem/ Science Denialism

Selected Publications

Talbot, C. J., Bennett, E. M., Cassell, K., Hanes, D. M., Minor, E. C., Paerl, H., . . . Xenopoulos, M. A. (2018). The impact of flooding on aquatic ecosystem services. Biogeochemistry, 141(3), 439-461. doi:10.1007/s10533-018-0449-7

Wollheim, W. M., Bernal, S., Burns, D. A., Czuba, J. A., Driscoll, C. T., Hansen, A. T., . . . Wohl, E. (2018). River network saturation concept: factors influencing the balance of biogeochemical supply and demand of river networks. Biogeochemistry, 141(3), 503-521. doi:10.1007/s10533-018-0488-0

Li, W., Zhai, L., Lei, Q., Wollheim, W. M., Liu, J., Liu, H., . . . Liu, S. (2018). Influences of agricultural land use composition and distribution on nitrogen export from a subtropical watershed in China. Science of The Total Environment, 642, 21-32. doi:10.1016/j.scitotenv.2018.06.048

Hopkinson, C. S., Morris, J. T., Fagherazzi, S., Wollheim, W. M., & Raymond, P. A. (2018). Lateral Marsh Edge Erosion as a Source of Sediments for Vertical Marsh Accretion. Journal of Geophysical Research: Biogeosciences, 123(8), 2444-2465. doi:10.1029/2017jg004358

Zuidema, S., Wollheim, W. M., Mineau, M. M., Green, M. B., & Stewart, R. J. (2018). Controls of Chloride Loading and Impairment at the River Network Scale in New England. Journal of Environmental Quality, 47(4), 839-847. doi:10.2134/jeq2017.11.0418

Deegan, L. A., Johnson, D. S., Warren, R. S., Peterson, B. J., Fleeger, J. W., Fagherazzi, S., & Wollheim, W. M. (2012). Coastal eutrophication as a driver of salt marsh loss. Nature, 490(7420), 388-392. doi:10.1038/nature11533

Beaulieu, J. J., Tank, J. L., Hamilton, S. K., Wollheim, W. M., Hall, R. O., Mulholland, P. J., . . . Thomas, S. M. (2011). Nitrous oxide emission from denitrification in stream and river networks. Proceedings of the National Academy of Sciences, 108(1), 214-219. doi:10.1073/pnas.1011464108

Harrison, J. A., Maranger, R. J., Alexander, R. B., Giblin, A. E., Jacinthe, P. -A., Mayorga, E., . . . Wollheim, W. M. (2009). The regional and global significance of nitrogen removal in lakes and reservoirs. Biogeochemistry, 93(1-2), 143-157. doi:10.1007/s10533-008-9272-x

Peterson, B. J. (2001). Control of Nitrogen Export from Watersheds by Headwater Streams. Science, 292(5514), 86-90. doi:10.1126/science.1056874

Mulholland, P. J., Tank, J. L., Sanzone, D. M., Wollheim, W. M., Peterson, B. J., Webster, J. R., & Meyer, J. L. (2000). NITROGEN CYCLING IN A FOREST STREAM DETERMINED BY A15N TRACER ADDITION. Ecological Monographs, 70(3), 471-493. doi:10.1890/0012-9615(2000)070[0471:nciafs]2.0.co;2

Most Cited Publications