Wilfred Wollheim

Co-director, Institute for the Study of Earth Oceans and Space
Phone: (603) 862-5022
Office: Complex Systems Research Center, James Hall Rm 276, Durham, NH 03824

In understanding the environment as an integrated system, WSAG explores the physical, chemical and biological processes that shape hydrological systems, with emphasis on the unique role of humans as agents of change. Our research and educational programs cross the boundaries of traditional scientific disciplines to foster interdisciplinary understanding of environmental change. Through field monitoring, whole ecosystem experiments, novel in situ sensor deployments, and dynamic hydrological and ecosystem modeling, we attempt to understand hydrological and biogeochemical dynamics at scales ranging from individual ecosystems, to whole river systems, to the global systems of inland waters. Built on strong collaborations that are national and international in scope, our group also serves as a teaching platform for the next generation of ecosystem and earth system scientists.

The success of the Water Systems Analysis Group reflects a rapidly emerging scientific field which pursues integrative studies of hydrology, biogeochemistry, and human-water interactions that is necessary to analyze the full dimension of anthropogenic change at local, regional, and global scales.


  • Ph.D., Earth Sciences, University of New Hampshire
  • M.S., Zoology/Physiology, University of Wyoming
  • B.S., Natural Resources, Cornell University

Courses Taught

  • INCO 790: Adv Rsrch Exp/Natural Resource
  • NR 504: Freshwater Resources
  • NR 707: Environmental Modeling
  • NR 751: Aquatic Ecosystems
  • NR 791: Proj/Environmental Science I
  • NR 905: Grant Writing
  • NR 993: Sem/ Science Denialism

Selected Publications

Wollheim, W. M., Bernal, S., Burns, D. A., Czuba, J. A., Driscoll, C. T., Hansen, A. T., . . . Wohl, E. (2018). River network saturation concept: factors influencing the balance of biogeochemical supply and demand of river networks. Biogeochemistry, 141(3), 503-521. doi:10.1007/s10533-018-0488-0

Hopkinson, C. S., Morris, J. T., Fagherazzi, S., Wollheim, W. M., & Raymond, P. A. (2018). Lateral Marsh Edge Erosion as a Source of Sediments for Vertical Marsh Accretion. Journal of Geophysical Research: Biogeosciences, 123(8), 2444-2465. doi:10.1029/2017jg004358

Koenig, L. E., Song, C., Wollheim, W. M., Rüegg, J., & McDowell, W. H. (2017). Nitrification increases nitrogen export from a tropical river network. Freshwater Science, 36(4), 698-712. doi:10.1086/694906

Wollheim, W. M., Mulukutla, G. K., Cook, C., & Carey, R. O. (2017). Aquatic Nitrate Retention at River Network Scales Across Flow Conditions Determined Using Nested In Situ Sensors. Water Resources Research, 53(11), 9740-9756. doi:10.1002/2017wr020644

Contosta, A. R., Adolph, A., Burchsted, D., Burakowski, E., Green, M., Guerra, D., . . . Wollheim, W. (2017). A longer vernal window: the role of winter coldness and snowpack in driving spring transitions and lags. Global Change Biology, 23(4), 1610-1625. doi:10.1111/gcb.13517

Deegan, L. A., Johnson, D. S., Warren, R. S., Peterson, B. J., Fleeger, J. W., Fagherazzi, S., & Wollheim, W. M. (2012). Coastal eutrophication as a driver of salt marsh loss. Nature, 490(7420), 388-392. doi:10.1038/nature11533

Beaulieu, J. J., Tank, J. L., Hamilton, S. K., Wollheim, W. M., Hall, R. O., Mulholland, P. J., . . . Thomas, S. M. (2011). Nitrous oxide emission from denitrification in stream and river networks. Proceedings of the National Academy of Sciences, 108(1), 214-219. doi:10.1073/pnas.1011464108

Webster, J. R., Mulholland, P. J., Tank, J. L., Valett, H. M., Dodds, W. K., Peterson, B. J., . . . Wollheim, W. M. (2003). Factors affecting ammonium uptake in streams - an inter-biome perspective. Freshwater Biology, 48(8), 1329-1352. doi:10.1046/j.1365-2427.2003.01094.x

Peterson, B. J. (2001). Control of Nitrogen Export from Watersheds by Headwater Streams. Science, 292(5514), 86-90. doi:10.1126/science.1056874

Mulholland, P. J., Tank, J. L., Sanzone, D. M., Wollheim, W. M., Peterson, B. J., Webster, J. R., & Meyer, J. L. (2000). NITROGEN CYCLING IN A FOREST STREAM DETERMINED BY A15N TRACER ADDITION. Ecological Monographs, 70(3), 471-493. doi:10.1890/0012-9615(2000)070[0471:nciafs]2.0.co;2

Most Cited Publications