Wilfred M. Wollheim

Associate Professor
Phone: (603) 862-5022
Office: Complex Systems Research Center, James Hall Rm 276, Durham, NH 03824


  • Ph.D., Earth Sciences, University of New Hampshire
  • M.S., Zoology/Physiology, University of Wyoming
  • B.S., Natural Resources, Cornell University

Courses Taught

  • INCO 790: Adv Rsrch Exp/Natural Resource
  • NR 707: Environmental Modeling
  • NR 751: Aquatic Ecosystems
  • NR 791: Proj/Environmental Science I
  • NR 795: Inv/Sediment Control Fluxes
  • NR 905: Grant Writing
  • NR 993: Sem/ Science Denialism
  • NR 995: Inv/DevOutreachWaterQuality

Selected Publications

Koenig, L. E., Song, C., Wollheim, W. M., Rüegg, J., & McDowell, W. H. (2017). Nitrification increases nitrogen export from a tropical river network. Freshwater Science, 36(4), 698-712. doi:10.1086/694906

Wollheim, W. M., Mulukutla, G. K., Cook, C., & Carey, R. O. (2017). Aquatic Nitrate Retention at River Network Scales Across Flow Conditions Determined Using Nested In Situ Sensors. Water Resources Research, 53(11), 9740-9756. doi:10.1002/2017WR020644

Contosta, A. R., Adolph, A., Burchsted, D., Burakowski, E., Green, M., Guerra, D., . . . Wollheim, W. (2017). A longer vernal window: the role of winter coldness and snowpack in driving spring transitions and lags. Global Change Biology, 23(4), 1610-1625. doi:10.1111/gcb.13517

Wlostowski, A. N., Gooseff, M. N., Bowden, W. B., & Wollheim, W. M. (2017). Stream tracer breakthrough curve decomposition into mass fractions: A simple framework to analyze and compare conservative solute transport processes. Limnology and Oceanography: Methods, 15(2), 140-153. doi:10.1002/lom3.10148

Samal, N. R., Wollheim, W. M., Zuidema, S., Stewart, R. J., Zhou, Z., Mineau, M. M., . . . Huber, M. (2017). A coupled terrestrial and aquatic biogeophysical model of the Upper Merrimack River watershed, New Hampshire, to inform ecosystem services evaluation and management under climate and land-cover change. Ecology and Society, 22(4). doi:10.5751/ES-09662-220418

Deegan, L. A., Johnson, D. S., Warren, R. S., Peterson, B. J., Fleeger, J. W., Fagherazzi, S., & Wollheim, W. M. (2012). Coastal eutrophication as a driver of salt marsh loss. Nature, 490(7420), 388-392. doi:10.1038/nature11533

Beaulieu, J. J., Tank, J. L., Hamilton, S. K., Wollheim, W. M., Hall, R. O., Mulholland, P. J., . . . Thomas, S. M. (2011). Nitrous oxide emission from denitrification in stream and river networks. Proceedings of the National Academy of Sciences, 108(1), 214-219. doi:10.1073/pnas.1011464108

Webster, J. R., Mulholland, P. J., Tank, J. L., Valett, H. M., Dodds, W. K., Peterson, B. J., . . . Wollheim, W. M. (2003). Factors affecting ammonium uptake in streams - an inter-biome perspective. Freshwater Biology, 48(8), 1329-1352. doi:10.1046/j.1365-2427.2003.01094.x

Peterson, B. J. (2001). Control of Nitrogen Export from Watersheds by Headwater Streams. Science, 292(5514), 86-90. doi:10.1126/science.1056874

Mulholland, P. J., Tank, J. L., Sanzone, D. M., Wollheim, W. M., Peterson, B. J., Webster, J. R., & Meyer, J. L. (2000). NITROGEN CYCLING IN A FOREST STREAM DETERMINED BY A 15 N TRACER ADDITION. Ecological Monographs, 70(3), 471-493. doi:10.1890/0012-9615(2000)070[0471:NCIAFS]2.0.CO;2

Most Cited Publications